Professor John Walsh

Emeritus Professor of Plant-Virus Interactions

Email: John.Walsh@warwick.ac.uk

Phone: 024 765 75028

Office (W): PPB 1.26

Office (GH): D 126

Walsh webpage

Research Clusters

Plant & Agricultural Biosciences

Microbiology & Infectious disease

Environment & Ecology

Warwick Centres and GRPs

Warwick Crop Centre

Vacancies and Opportunities

For PhD and postdoctoral opportunities, and interest in potential collaborations, please contact me at the above email address.

Research Interests

Plants, like most living things, are infected by viruses. These viruses can cause major diseases and are a major constraint limiting crop production worldwide, but particularly in underdeveloped regions.

Withdrawal of pesticide active ingredients is exacerbating problems Plant viruses are responsible for 47% of new/emerging infectious diseases of plants.

Current Research and Teaching Interests:

Our group has a diverse research programme ranging from fundamental research through translational to very applied research directly relevant to industry and farmers. These include:

Plant - virus interactions
Identifying, characterising and understanding the mechanism of plant resistance genes with a view to developing durable disease resistance
Studying plant virus diversity
Exploring the insect vectors of plant viruses
Investigating viruses in wild plants
Researching plant viruses transmitted by fungi and single-celled organisms

Research: Technical Summary

Our group has a diverse research programme ranging from fundamental research through translational to very applied research directly relevant to growers. These include:

Plant - virus interactions, particularly viruses infecting brassicas (turnip mosaic virus, turnip yellows virus etc.)
Identifying, characterising, mapping and understanding the mechanism of plant resistance genes and identifying their viral determinants with a view to developing durable plant resistances
Understanding virus - virus interactions in plants and viral fitness
Plant virus diversity
Plant - virus co-evolution in wild plants
Plant viruses transmitted by fungi and protists and the vectors themselves

John's group have a number of patents and have had a number of products commercialised including germplasm possessing resistance genes to plant viruses and diagnostic reagents for detection of plant viruses and protists.

Biography

John studied Applied Biology at the University of Leeds and then went on to a co-operative award PhD at Rothamsted Experimental Station (now Rothamsted Research) and University of Leeds in Plant Pathology. During his PhD studies, John was awarded the Best Student Paper Award, from the Society of Nematologists at their meeting in Salt Lake City. This was followed by a Potato Marketing Board postdoctoral fellowship at Rothamsted. John arrived at the Wellesbourne Crop Centre site in 1983 and is currently group leader for the Plant - Virus Interactions group. This group has a broad range of research projects from Applied, through Strategic to Fundamental, funded from a wide variety of sources. The underlying aim is to provide environmentally friendly sustainable disease control measures whilst at the same time reducing inputs.

John has served his field as a member of the Association of Applied Biologists Virology Group Committee, secretary and president of to the International Working Group on Legume and Vegetable Viruses, Microbilogy Society Virology Division Committee and NERC Peer Review College. He has also been an editor of a number of journals including including the Annals of Applied Biology, the European Journal of Plant Pathology, General Plant Pathology and Horticultural Plant Journal.

Publications

Beddoe, Naomi, McGreig, Sam, Walsh, John A., Fox, Adrian, Adams, Ian P. and Forsyth, Alec (2024) Nerine potexvirus 1 : a new potexvirus species detected from nerine in the United Kingdom. Journal of Plant Pathology, 106 . pp. 403-408. doi:10.1007/s42161-023-01512-z ISSN 1125-4653.

Pimenta, Ricardo J. G., MacLeod, Kyle, Babb, Robyn, Coleman, Kaitlyn, MacDonald, Joni, Asare-Bediako, Elvis, Newbert, Max John, Jenner, Carol E. and Walsh, John A. (2024) Genetic variation of turnip yellows virus in arable and vegetable brassica crops, perennial wild brassicas and aphid vectors collected from the plants. Plant Disease, 108 (3). pp. 616-623. doi:10.1094/pdis-05-23-0906-re ISSN 1943-7692.

MacLeod, Kyle, Greer, Shannon F., Bramham, Lawrence, Pimenta, Ricardo J. G., Nellist, Charlotte F., Hackenburg, Dieter, Teakle, Graham R., Barker, Guy C. and Walsh, John A. (2023) A review of sources of resistance to turnip yellows virus ( TuYV ) in Brassica species. Annals of Applied Biology, 183 (3). pp. 200-208. doi:10.1111/aab.12842 ISSN 0003-4746.

Walsh, Hanna M., Rönkä, Antti and Walsh, John A. (2023) Identification and genetic inheritance of a new source of broad-spectrum extreme resistance to turnip mosaic virus (TuMV) in Brassica rapa. European Journal of Plant Pathology, 165 . pp. 693-699. doi:10.1007/s10658-022-02634-3 ISSN 0929-1873.

Nellist, Charlotte F., Ohshima, Kazusato, Ponz, Fernando and Walsh, John A. (2022) Turnip mosaic virus, a virus for all seasons. Annals of Applied Biology, 180 (3). pp. 312-327. doi:10.1111/aab.12755 ISSN 0003-4746.

Bramham, Lawrence E., Wang, Tongtong, Higgins, Erin E., Parkin, Isobel A. P., Barker, Guy C. and Walsh, John A. (2022) Characterization and mapping of retr04, retr05 and retr06 broad-spectrum resistances to Turnip mosaic virus in Brassica juncea, and the development of robust methods for utilizing recalcitrant genotyping data. Frontiers in Plant Science, 12 . doi:10.3389/fpls.2021.787354 ISSN 1664-462X.

Greer, Shannon F., Hackenberg, Dieter, Gegas, Vasilis, Mitrousia, Georgia, Edwards, David, Batley, Jacqueline, Teakle, Graham R., Barker, Guy C. and Walsh, John A. (2021) Quantitative trait locus mapping of resistance to turnip yellows virus in Brassica rapa and Brassica oleracea and introgression of these resistances by resynthesis into allotetraploid plants for deployment in Brassica napus. Frontiers in Plant Science, 12 . 781385. doi:10.3389/fpls.2021.781385 ISSN 1664-462X.

Greer, S. F., Newbert, M. J., Rodrigues, L. K., Oliveira, A. M., Chaves, A. L. R., Calegario, R. F., Barker, G. C., Eiras, M. and Walsh, John A. (2021) First report of Turnip yellows virus in Brazil. New Disease Reports, 44 (2). e12045. doi:10.1002/ndr2.12045 ISSN 2044-0588.

Rodrigues, Leilane Karam, Chaves, Alexandre Levi Rodrigues, Kitajima, Elliot Watanabe, Calegario, Renata Faier, Brunelli, Katia Regiane, da Silva, Fabio Nascimento, Harakava, Ricardo, Walsh, John A. and Eiras, Marcelo (2021) Characterisation of turnip mosaic virus isolates reveals high genetic variability and occurrence of pathotype 1 in Brazil. European Journal of Plant Pathology, 160 . pp. 883-900. doi:10.1007/s10658-021-02291-y ISSN 0929-1873.

MacLeod, Kyle, Greer, Shannon F., Hackenberg, Dieter, Bramham, Lawrence, Newbert, Max John, Hambidge, A. J. (Angela J.), Katschnig, Diana, Baker, Adam R., Teakle, Graham R., Barker, Guy C. and Walsh, John A. (2021) New sources of resistance to turnip yellows virus from Brassica oleracea and Brassica rapa and their introgression into oilseed rape. In: Management of Diseases and Pests of Oilseed Rape, University of Hertfordshire, 16 Jun 2021. Published in: Proceedings of Management of Diseases and Pests of Oilseed Rape, University of Hertfordshire, 16 June 2021 pp. 55-58.

Hammond, J., Adams, I., Fowkes, A., McGreig, S., Botermans, M., van Oorspronk, J., Westenberg, M., Verbeek, M., Dullemans, A., Stijger, C., Blouin, A., Massart, S., de Jonghe, K., Heyneman, M., Walsh, John A. and Fox, A. (2021) Sequence analysis of 43-year old samples of Plantago lanceolata show that Plantain virus X is synonymous with Actinidia virus X and is widely distributed. Plant Pathology, 70 (2). pp. 249-258. doi:10.1111/ppa.13310 ISSN 0032-0862.

Asare‐Bediako, Elvis, Jones, Julie E. , Hambidge, Angela, Stevenson, Mark, Mead, Andrew, Jenner, Carol E. and Walsh, John A. (2020) The incidence of Turnip yellows virus in oilseed rape crops (Brassica napus L.) in three different regions of England over three consecutive growing seasons and the relationship with the abundance of flying Myzus persicae. Annals of Applied Biology, 176 (2). pp. 130-137. doi:10.1111/aab.12565 ISSN 0003-4746.

Hackenberg, Dieter, Asare-Bediako, Elvis, Baker, Adam R., Walley, Peter Glen, Jenner, Carol E., Greer, Shannon, Bramham, Lawrence, Batley, Jacqueline, Edwards, David, Delourme, Regine, Barker, Guy C., Teakle, Graham R. and Walsh, John A. (2020) Identification and QTL mapping of resistance to Turnip yellows virus (TuYV) in oilseed rape, Brassica napus. Theoretical and Applied Genetics, 133 (2). pp. 383-393. doi:10.1007/s00122-019-03469-z ISSN 0040-5752.

Kapoor, Shelly, Handa, Anil, Walsh, John A. and Sharma, Rajnish (2020) Confirmation of Radish Isolate of Turnip mosaic virus in India through biological and serological evidences. Plant Pathology Journal, 19 (4). pp. 211-220. doi:10.3923/ppj.2020.211.220 ISSN 1812-5387.

Michelmore, Richard , Coaker, Gitta , Bart, Rebecca , Beattie, Gwyn , Bent, Andrew , Bruce, Toby , Cameron, Duncan , Dangl, Jeff , Dinesh-Kumar, Savithramma , Edwards, Rob et al.

(2017) Foundational and translational research opportunities to improve plant health. Molecular Plant-Microbe Interactions, 30 (7). pp. 515-516. doi:10.1094/MPMI-01-17-0010-CR ISSN 0894-0282.

Jannat, Shopan, Mou, Haipeng, Zhang, Lili, Zhang, Changtong, Ma, Weiwei, Walsh, John A., Hu, Zhongyuan, Yang, Jinghua and Zhang, Mingfang (2017) Eukaryotic translation initiation factor 2B-beta (eIF2B β), a new class of plant virus resistance gene. The Plant Journal, 90 (5). pp. 929-940. doi:10.1111/tpj.13519 ISSN 0960-7412.

Afonso, Claudio L., Amarasinghe, Gaya K., Bányai, Krisztián, Bào, Yīmíng, Basler, Christopher F., Bavari, Sina, Bejerman, Nicolás, Blasdell, Kim R., Briand, François-Xavier, Briese, Thomas et al.

(2016) Taxonomy of the order Mononegavirales : update 2016. Archives of Virology, 161 (8). pp. 2351-2360. doi:10.1007/s00705-016-2880-1 ISSN 0304-8608.

Sánchez, Flora, Manrique, Pilar, Mansilla, Carmen, Lunello, Pablo, Wang, Xiaowu, Rodrigo, Guillermo, López-González, Silvia, Jenner, Carol E., González-Melendi, Pablo, Elena, Santiago F., Walsh, John A. and Ponz, Fernando (2015) Viral strain-specific differential alterations in Arabidopsis developmental patterns. Molecular Plant-Microbe Interactions, 28 (12). pp. 1304-1315. doi:10.1094/MPMI-05-15-0111-R ISSN 0894-0282.

Schwinghamer, Mark W., Schilg, Mark A., Walsh, John A., Bambach, Rodney W., Cossu, Rosa M., Bambridge, Judith M., Hind-Lanoiselet, Tamrika L., McCorkell, Bruce E. and Cross, Peter (2014) Turnip mosaic virus : potential for crop losses in the grain belt of New South Wales, Australia. Australasian Plant Pathology, 43 (6). pp. 663-678. doi:10.1007/s13313-014-0304-9 ISSN 0815-3191.

Lydiate, D. J., Rusholme, Rachel L., Higgins, E. E. and Walsh, John A. (2014) Genetic control of immunity to turnip mosaic virus (TuMV) pathotype 1 in Brassica rapa (Chinese cabbage). Genome , 57 (8). pp. 419-425. doi:10.1139/gen-2014-0070 ISSN 0831-2796.

Nellist, Charlotte F., Qian, Wei, Jenner, Carol E., Moore, Jonathan D., Zhang, Shujiang, Wang, Xiaowu, Briggs, William H., Barker, Guy C., Sun, Rifei and Walsh, John A. (2014) Multiple copies of eukaryotic translation initiation factors in Brassica rapafacilitate redundancy, enabling diversification through variation in splicing and broad-spectrum virus resistance. The Plant Journal, Volume 77 (Number 2). pp. 261-268. doi:10.1111/tpj.12389 ISSN 0960-7412.

Qian, Wei, Zhang, Shujiang, Zhang, Shifan, Li, Fei, Zhang, Hui, Wu, Jian, Wang, Xiaowu, Walsh, John A. and Sun, Rifei (2013) Mapping and candidate-gene screening of the novel Turnip mosaic virus resistance gene retr02 in Chinese cabbage (Brassica rapa L.). Theoretical and Applied Genetics, Volume 126 (Number 1). pp. 179-188. doi:10.1007/s00122-012-1972-x ISSN 0040-5752.

Nguyen, Huy D., Tomitaka, Yasuhiro, Ho, Simon Y. W., Duchêne, Sebastián, Vetten, Heinrich-Josef, Lesemann, D.-E., Walsh, John A., Gibbs, Adrian J. and Ohshima, Kazusato (2013) Turnip mosaic potyvirus probably first spread to Eurasian brassica crops from wild orchids about 1000 years ago. PLoS One, Volume 8 (Number 2). Article no.e55336. doi:10.1371/journal.pone.0055336 ISSN 1932-6203.

Walsh, J.A. and Verbeek, M. (2012) Genus – Varicosavirus. In: Virus Taxonomy — Ninth Report of the International Committee on Taxonomy of Viruses. International Committee on Taxonomy of Viruses, Vol.9 . Elsevier, pp. 777-781. ISBN 978-0123-84684-6

Jenner, Carol E., Nellist, Charlotte F., Barker, Guy C. and Walsh, John A. (2010) Turnip mosaic virus (TuMV) is able to use alleles of both eIF4E and eIF(iso)4E from multiple loci of the diploid Brassica rapa. Molecular Plant - Microbe Interactions, Vol.23 (No.11). pp. 1498-1505. doi:10.1094/MPMI-05-10-0104 ISSN 0894-0282.

Ho, Thien, Wang, Liang, Huang, Linfeng, Li, Zhigang, Pallett, Denise W., Dalmay, Tamas, Ohshima, Kazusato, Walsh, John A. and Wang, Hui (2010) Nucleotide bias of DCL and AGO in plant anti-virus gene silencing. Protein & Cell, Vol.1 (No.9). pp. 847-858. doi:10.1007/s13238-010-0100-4 ISSN 1674-8018 .

Hartwright, Lucy M., Hunter, Paul J. and Walsh, John A. (2009) A comparison of Olpidium isolates from a range of host plants using internal transcribed spacer sequence analysis and host range studies. Fungal Biology, Vol.114 (No.1). pp. 26-33. doi:10.1016/j.mycres.2009.09.008 ISSN 0953-7562.

Lewsey, Mathew, Surette, Monique, Robertson, Fiona C. , Ziebell, Heiko, Choi, Sun Hee , Ryu, Ki Hyun , Canto, Tomas, Palukaitis, Peter, Payne, Tina, Walsh, John A. and Carr, John P. (2009) The role of the Cucumber mosaic virus 2b protein in viral movement and symptom induction. Molecular Plant - Microbe Interactions, Vol.22 (No.6). pp. 642-654. doi:10.1094/MPMI-22-6-0642 ISSN 0894-0282.

Barker, Guy C., Jenner, Carol E. and Walsh, John A. (2009) Possible role of Brassica rapa translation initiation factors in broad-spectrum resistance to Turnip mosaic virus. In: An International Conference: Advances in Plant Virology, Harrogate International Centre, Harrogate, UK , 1-3 April 2009. Published in: Programme for An International Conference: Advances in Plant Virology 2009

Walsh, John A. (2009) Reducing losses from virus-induced storage disorders of processing cabbage. United Kingdom: Horticultural Development Company. (HDC News).

Asare-Bediako, Elvis, Jenner, Carol E., Stevens, M. and Walsh, John A. (2009) The incidence and diversity of Turnip yellows virus (TuYV) in oilseed rape (Brassica napus) in the UK. In: An International Conference: Advances in Plant Virology, Harrogate International Centre, Harrogate, UK , 1-3 April 2009. Published in: Programme for An International Conference: Advances in Plant Virology 2009

Haj Kassem, Amin A. and Walsh, John A. (2008) Characterising resistance to Turnip mosaic virus (TuMV) in Turnip (Brassica rapa rapa). Arab Journal of Plant Protection, Vol.26 (No.2). pp. 168-172. ISSN 0255-982X.

Walsh, John A. (2008) Devastating crop virus could be eradicated. [Online]. (http://www2.warwick.ac.uk/newsandevents/icast/arch...). Coventry: University of Warwick.

Sanchez, F., Lunello, P., Mansilla, C., Martínez, F., Wang, Xiaow, Walsh, John A., Jenner, Carol E. and Ponz, F. (2008) Disturbance of Arabidopsis thaliana development by a potyviral infection maps to the P3/p6k1 viral genomic region. In: 4th EPSO Conference , Toulon (Côte d’Azur), France , 22–26 June, 2008. Published in: Proceedings of the 4th EPSO Conference “Plants for Life" Article No. P 086 .

Walsh, John A., Jenner, Carol E., Bambridge, J.M., Rusholme, Rachel L., Hughes, S., Higgins, Erin E., Parkin, I.A.P, Barker, Guy C. and Lydiate, Derek J. (2008) Genetic Resistance to Turnip mosaic virus (TuMV) in Brassicas. In: Brassica 2008: 5th ISHS International Symposium on Brassicas and the 16th Crucifer Genetics Workshop, Lillehammer, Norway, 8 - 12 Sept, 2008. Published in: Preceedings of the 5th ISHS International Symposium on Brassicas and the 16th Crucifer Genetics Workshop (F21). p. 147.

Obermeier, C., Hunter, Paul J., Machado, R., Ohshima, Kazusato, Kearsey, M. J. and Walsh, John A. (2008) Plant – virus co-evolution in wild brassicas. In: 3rd Conference of the International Working Group on Legume and Vegetable Viruses (IWGLVV) , Ljubljana, Slovenia, 20th – 23rd Aug, 2008 . Published in: Proceedings of the 3rd Conference of the International Working Group on Legume and Vegetable Viruses (IWGLVV) p. 23.

Pallett, D. W., Cooper, J. I., Wang, H., Reeves, J., Luo, Z., Machado, R., Obermeier, C., Walsh, John A. and Kearsey, M. J. (2008) Variation in the pathogenicity of two Turnip mosaic virus isolates in wild UK Brassica rapa provenances. Plant Pathology, Vol.57 (No.3). pp. 401-407. doi:10.1111/j.1365-3059.2007.01764.x ISSN 0032-0862.

Carr, J.P., Lewsey, Mathew, Westwood, J., Ziebell, Heiko, MacLean, Daniel, Kleczkowski, A., Stevens, M. and Walsh, John A. (2008) The effects of cucumber mosaic virus 2b protein on methyl jasmonate induced transcriptional responses and plant-aphid interactions in Arabidopsis. In: American Society for Virology 27th Annual Meeting, Cornell University, Ithaca, NY, Jul 12-16, 2008. Published in: Proceedings of the 27th Annual Meeting of American Society for Virology

Rusholme, Rachel L., Higgins, Erin E., Walsh, John A. and Lydiate, Derek J. (2007) Broad-spectrum resistance to turnip mosaic virus in Brassica rapa (Chinese cabbage). In: International Conference, Genetic control of plant pathogenic viruses and their vectors: Towards new resistance strategies, El Puerto de Santa Maria, Cadiz, Spain, 23-27 Nov 2008. Published in: Proceedings of the Genetic control of plant pathogenic viruses and their vectors: Towards new resistance strategies doi:10.1099/vir.0.83194-0

Rusholme, Rachel L., Higgins, Erin E., Walsh, John A. and Lydiate, Derek J. (2007) Genetic control of broad-spectrum resistance to turnip mosaic virus in Brassica rapa (Chinese cabbage). Journal of General Virology, Vol.88 (Part 11). pp. 3177-3186. doi:10.1099/vir.0.83194-0 ISSN 0022-1317.

Walsh, John A. Avoid the slow burn in store. HDC News, Vol.149 . pp. 18-19.

Walsh, John A. Brassica and Lettuce Propagation: Identifying means of killing clubroot and Olpidium resting spores in trays to avoid infection of transplants. United Kingdom: Horticultural Development Company. (HDC News).

Jenner, Carol E., Barker, Guy C., Walsh, John A., Hoffinger, B., Hammond-Kosack, K., Beaudoin, F. and Kanyuka, K. Exploiting eIF4E-based and associated broad-spectrum recessive resistance to potyviruses in dicots and monocots. In: 2nd BBSRC Innovation in Crop Science - exploitation of genetics for sustainability initiative grant holders workshop, Cirencester, England, 28-29 Jan 2009. Published in: Crop Science Initiative outcomes, November 2010

Research Projects

Title	Funder	Award start	Award end
MTA - DSMZ GmbH - polyclonal antibody against BWYV	Leibniz-Institut DSMZ	15 Jan 2020	14 Jan 2025
Horticulture CRISPR Cas9-mediated and inter-species transfer of disease resistance to crop plants	BBSRC	01 Jul 2023	22 Mar 2024
BBSRC FoF Collaboration Agreement with Syngenta, Limagrain, LS Plant Breeding and Elsoms (Limagrain Contribution) // Linked to IDEATE 61514 // R.LFJW.4018 - August 2022	Limagrain UK Ltd	11 Feb 2020	31 Dec 2023
BBSRC FoF Collaboration Agreement with Syngenta, Limagrain, LS Plant Breeding and Elsoms (Syngenta contribution) // Linked to IDEATE 61514 // R.LFJW.4018 - August 2022	Syngenta UK Ltd	11 Feb 2020	31 Dec 2023
h20 SMARTPROTECT: SMART agriculture for innovative vegetable crop PROTECTion: harnessing advanced methodologies and technologies	European Commission	01 Jan 2020	30 Jun 2023
Genomics-led improvement of biotic and abiotic stress tolerance in mustard rape for economic and environmental sustainability	BBSRC	01 Sep 2018	31 Aug 2022
CoA: Genomics-led improvement of biotic and abiotic stress tolerance in mustard rape for economic and environmental sustainability; linked to Ideate 56617	UK Research and Innovation	01 Sep 2021	30 Sep 2021
CoA: Delivering Important Virus Resistance (DIVR) - Follow on Fund; linked to Ideate 61514	UK Research and Innovation	01 May 2021	30 Sep 2021
CoA: Developing integrated approaches for pest and disease control in horticultural field crops (IAPAD); ; linked to Ideate 45111	UK Research and Innovation	04 Jan 2021	30 Apr 2021
Deployment of durable, broad-spectrum resistance to Turnip mosaic virus into Brassica oleracea in sub-Saharan Africa	BBSRC	01 Jan 2019	31 Dec 2019
Developing integrated approaches for pest and disease control in horticultural field crops (IAPAD)	BBSRC	01 Apr 2015	23 Oct 2019
Exploiting new sources of broad-spectrum resistance to Turnip mosaic virus in brassicas (iCASE)	Sakata UK Ltd	05 Oct 2015	04 Oct 2019
Vegetable Genetic Improvement Network (VeGIN) extension 2017	Department for Environment, Food and Rural Affairs	01 Apr 2017	31 Mar 2018
CIRC training grant	BBSRC	01 Oct 2011	30 Sep 2015
Support towards 4 year BBSRC CIRC studentship (Max Newbert) travel/conferences only	Perry Foundation	01 Oct 2011	30 Sep 2015
Exploiting sources of resistance to turnip yellow virus for deployment in oilseed - CIRC proposal	BBSRC	01 Sep 2011	31 Aug 2015
The incidence of turnip yellow virus (TuYV) in over wintered cauliflower and brussel sprouts	AHDB (Formerly HDC)	01 May 2010	30 Apr 2013
Brassica and lettuce propagation: Scaling up means of controlling clubroot resting spores in trays to avoid infection of transplants.	AHDB (Formerly HDC)	01 Apr 2011	31 Jan 2013
Amendment to RLFJW0502 (26890) The time of infection of overwintered cauliflower and Brussels sprout by Turnip yellows virus (TuYV) and the potential of insecticides to control the virus (FV365a)	AHDB (Formerly HDC)	01 Apr 2011	31 May 2012
IF0157 Vegetable Genetic Improvement Network (VeGIN): Leafy vegetables and salads	Department for Environment, Food and Rural Affairs	01 Apr 2009	31 Mar 2012
Introgression of broad-spectrum resistance to turnip mosaic virus (Tumv) in to Brassica rapa	Syngenta Ltd	01 Jan 2009	31 Dec 2011
Studentship: Deploying virus resistance in brassicas and understanding the interaction between the viral VPg protein and brassica eIF4E and eIF(iso)4E proteins (CASE)	Syngenta Ltd	01 Oct 2008	30 Sep 2011
FV0160b / 13540 Reducing losses from Virus-induced storage disorders of processing cabbage	Horticultural Development Council	01 Feb 2006	30 Jun 2010
Brassica & Lettuce propagation: Identifying means of killing clubroot and olipidium resting spores	Horticultural Development Council	01 Apr 2008	17 May 2010
Exploiting eIF4E - based and associated broad spectrum recessive resistance to potyviruses in dicots and monocots.	BBSRC	01 Apr 2007	31 Mar 2010
Wild brassica screen for virus resistance (Wibsvir). Pre-breeding research to support sutainable farming of brassicas in the UK	Department for Environment, Food and Rural Affairs	01 Jan 2009	31 Jan 2010
IF0191 Wild brassica screen for virus resistance (wibsvir). Pre-breeding research to support farming of brassicas in the UK	Department for Environment, Food and Rural Affairs	01 Jan 2009	30 Sep 2009
Crop Improvement of Field Vegetables	Department for Environment, Food and Rural Affairs	01 Apr 2004	31 Mar 2009
Watercress: Optimising Zinc applications for controlling crook root disease and minimising zin discharges.	Horticultural Development Council	01 Jan 2003	01 Sep 2004

Teaching / Student Supervision